In 1858 Charles Darwin was forty-nine years old, a gentleman scientist living quietly at Down House in the Kent countryside, respected by fellow biologists and well liked among his wide and distinguished circle of acquaintances. He was not yet a focus of debate; his “big book on species” still lay on his study desk in the form of a huge pile of manuscript. For more than twenty years he had been accumulating material for it, puzzling over questions it raised, trying—it seemed endlessly—to bring it to a satisfactory conclusion. Publication appeared to be as far away as ever, delayed by his inherent cautiousness and wish to be certain that his startling theory of evolution was correct. It is at this point that the concluding volume of Janet Browne’s biography opens. The much-praised first volume, Voyaging, carried Darwin’s story through his youth and scientific apprenticeship, the adventurous Beagle voyage, his marriage and the birth of his children, the genesis and development of his ideas. Now, beginning with the extraordinary events that finally forced the Origin of Species into print, we come to the years of fame and controversy. For Charles Darwin, the intellectual upheaval touched off by his book had deep personal as well as public consequences. Always an intensely private man, he suddenly found himself and his ideas being discussed—and often attacked—in circles far beyond those of his familiar scientific community. Demonized by some, defended by others (including such brilliant supporters as Thomas Henry Huxley and Joseph Hooker), he soon emerged as one of the leading thinkers of the Victorian era, a man whose theories played a major role in shaping the modern world. Yet, in spite of the enormous new pressures, he clung firmly, sometimes painfully, to the quiet things that had always meant the most to him—his family, his research, his network of correspondents, his peaceful life at Down House. In her account of this second half of Darwin’s life, Janet Browne doe